Security Type

Security Class

Title

Fee

Calculation

Rule

Amount
Registered

Proposed
Maximum
Offering
Price Per
Unit

Maximum
Aggregate
Offering

Price

Fee

Rate

Amount of
Registration
Fee

Equity

Common Stock, par value $0.001 per share, reserved for issuance under the Registrant's 2014 Equity Incentive Plan

Rule 457(c) and Rule 457(h)

895,346 (2)

$23.24(4)

$20,807,841.04

0.00011020

$2,293.03

Equity

Common Stock, par value $0.001 per share, reserved for issuance under the Registrant's 2014 Employee Stock Purchase Plan

Rule 457(c) and Rule 457(h)

179,069 (3)

$19.76(5)

$3,538,403.44

0.00011020

$389.94

Total Offering Amounts

1,074,415

$24,346,244.48

Total Fee Offsets(6)

—

Net Fee Due

$2,682.97

(1)

Pursuant to Rule 416(a) of the Securities Act of 1933, as amended (the “Securities Act”), this registration statement (the “Registration Statement”) shall also cover any additional shares of the Registrant’s Common Stock that become issuable under the Registrant’s 2014 Equity Incentive Plan (the “2014 Plan”) or the Registrant’s 2014 Employee Stock Purchase Plan (the “2014 ESPP”) by reason of any stock dividend, stock split, recapitalization or other similar transaction effected without receipt of consideration or conversion of the Registrant’s Common Stock that increases the number of outstanding shares of Common Stock.

(2)

Represents an automatic increase in the number of shares of Common Stock reserved for issuance pursuant to future awards under the 2014 Plan as a result of the annual evergreen increase under the 2014 Plan.

(3)

Represents an automatic increase in the number of shares of Common Stock reserved for issuance pursuant to future awards under the 2014 ESPP as a result of the annual evergreen increase under the 2014 ESPP.

(4)

Pursuant to Rule 457(c) and 457(h) under the Securities Act, and solely for the purpose of calculating the registration fee, the proposed maximum offering price per share is $23.24, which is the average of the high and low prices of shares of Common Stock on The Nasdaq Global Select Market (“Nasdaq”) on February 23, 2023 (such date being within five business days of the date that this Registration Statement was filed with the U.S. Securities and Exchange Commission (the “SEC”)).

(5)

Pursuant to Rule 457(c) and 457(h) under the Securities Act, and solely for the purpose of calculating the registration fee, the proposed maximum offering price per share is $19.76, which is 85% of the average of the high and low prices of shares of Common Stock on Nasdaq on Febraury 23, 2023 (such date being within five business days of the date that this Registration Statement was filed with the SEC). Pursuant to the 2014 ESPP, the purchase price of the shares of common stock reserved for issuance thereunder will be 85% of the lower of the fair market value of the common stock on (i) the first trading day of the offering period or (ii) the exercise date.

(6)

The Registrant does not have any fee offsets.